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Winners of the 2016 National Geographic Nature Photographer of the Year Contest (18 photos)

National Geographic has announced the winners of its annual photo contest, with the Grand Prize Winner Greg Lecoeur receiving a 10-day trip for two to the Galapagos Islands. National Geographic was once again kind enough to let us display the winning images here. The captions below were written by the individual photographers.

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Sardine Run, Grand Prize Winner, and First Place Action Winner. During the sardine migration along the Wild Coast of South Africa, millions of sardines are preyed upon by marine predators such as dolphins, marine birds, sharks, whales, penguins, sailfishes, and sea lions. The hunt begins with common dolphins that have developed special hunting techniques to create and drive bait balls to the surface. In recent years, probably due to overfishing and climate change, the annual sardine run has become more and more unpredictable. It took me two weeks to have the opportunity to witness and capture this marine predation. (© Copyright G. Lecoeur / 2016 National Geographic Nature Photographer of the Year)
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http://www.theatlantic.com/photo/2016/12/winners-of-the-2016-national-geographic-nature-photographer-of-the-year-contest/510350/